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Astyanax dnophos, new species, is described from the lower rio Xingu, Pará, Brazil. The new species is distinguished from its purported congeners by the presence of a broad longitudinal, midlateral dark stripe that extends from the snout to the caudal peduncle. Astyanax dnophos dwells on shadowed, rocky shelters on rapids, and it is apparently one of the few species of Characidae outside Serrasalminae reported to be rheophilic.