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The synergidless female gametophyte of Plumbago zeylanica receives the pollen tube through specialized cell wall ingrowths at the base of the egg; tube growth continues between egg and central cells. Pollen tube discharge occurs between egg and central cell and results in release of two male gametes, vegetative nucleus, and some pollen cytoplasm. Except for the location of gamete discharge, details of transmission and fusion of gametic nuclei appear to conform to reports of these processes in taxa possessing conventional embryo sacs.