Genetic dissection of the grain-filling rate and related traits through linkage analysis and genome-wide association study in bread wheat oleh Hai-xia YU, Xi-xian DUAN, Ai-qing SUN, Xiao-xiao SUN, Jing-juan ZHANG, Hua-qing SUN, Yan-yan SUN, Tang-yuan NING, Ji-chun TIAN, Dong-xue WANG, Hao LI, Ke-xin FAN, Ai-ping WANG, Wu-jun MA, Jian-sheng CHEN